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These countries, located near the geographical extremes of Europe, were chosen to capture a large fraction of European phenotypic variation. For eye pigmentation, the iris color score C was calculated from the analysis of digital photographs taken under controlled light conditions using a cardboard box with form fitting foam padding such that the only illumination was from the camera flash. Studies including larger sample sizes are needed to characterize the RQ variant effect on skin pigmentation. View Article Google Scholar 3. The LCT gene is a locus well-documented as showing substantial genetic differentiation within Europe [29] , [30].
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